He then started his research career at the University of St Andrews by investigating cells in the Temporal cortex with David Perrett, and described cells that respond when the monkey views particular faces in a way that correlates with conscious perception.
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As a student he was a member of the Studienstiftung, Germany's most prestigious foundation for the promotion of excellence in study.
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A conceptually similar model proposed by Christian Keysers and David Perrett, based on what we know about the neural properties of mirror neurons and Spike-timing-dependent plasticity is the Hebbian Learning Account of Mirror Neurons.