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He reevaluates August Weismann's model of the cell compartmentalization of somatic and germline cell lineages (see Weismann barrier), and argues that the vision of the individual taken by the modern synthesis is insufficient to explain the early evolution of development or ontogeny.
She took a first degree in genetics Cambridge University, and DPhil Oxford University in early gene mapping studies using somatic cell hybrids, with Walter Bodmer.