In compiling this, the authors used a variety of phylogenetic information including molecular analysis, anatomical analysis, shell morphology and shell microstructure as well as bio-geographic, paleobiogeographic and stratigraphic information.
Preliminary phylogenetic studies in 2006 using molecular and morphological evidence have suggested that the genus forms three natural species confined to the southeastern United States.
The Korean hare (Lepus coreanus) was at one time considered to be a subspecies of the Chinese hare but molecular studies of mtDNA have since shown that the Korean hare is in fact a separate species.
Molecular phylogenetics has revealed that this species also includes the animals formerly separated as C. gymnurus, C. neglectus, C. tylorhinus and C. zinseri.
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A study of the molecular phylogeny of bishops and widowbirds published in 2008 found that it formed part of a clade along with the Fire-fronted Bishop (E. diadematus), Black-winged Red Bishop (E. hordeaceus), Northern Red Bishop (E. franciscanus), Southern Red Bishop (E. orix), Zanzibar Red Bishop (E. nigroventris) and Red-collared Widowbird (E. ardens).
In 1999, under the new joint editorship of Martin Kreitman (the current editor-in-chief), Takashi Gojobori, and Giorgio Bernardi, the journal made explicit its focus on "the fundamental issues in molecular evolution", to the exclusion of molecular phylogenetics (the purview of several other molecular evolution journals, most significantly Molecular Phylogenetics and Evolution) and origin of life research, except in cases that have "a special significance and impact".
So far, most of these groups have been defined cladistically rather than being defined as formal Linnean taxa, though there have been several cases in which older proposed segregates from the Tricholomataceae have been validated by evidence coming from molecular phylogenetics.
The authors placed the new species into Peter Taylor's section Psyllosperma, which has subsequently been merged with section Foliosa based on molecular phylogenetics.